Winter Wren

The winter wren was described and illustrated in 1808 by the American ornithologist Alexander Wilson. He was uncertain as to whether the wren should be considered a separate species or a subspecies of the Eurasian wren. When Louis Pierre Vieillot, a French ornithologist, described the winter wren in 1819 he considered it a separate species and coined the current binomial name Troglodytes hiemalis. The specific epithet is Latin and means "of winter". The type locality was restricted to Nova Scotia by Harry C. Oberholser in 1902. The winter wren was formerly considered to be conspecific with the Eurasian wren (Troglodytes troglodytes) and the Pacific wren (Troglodytes pacificus). The two North American species were split from the Eurasian wren based on a study of mitochondrial DNA published in 2007. A study published in 2008 of the songs and genetics of individuals in an overlap zone between Troglodytes hiemalis and Troglodytes pacificus found strong evidence of reproductive isolation between the two. It was suggested that the subspecies pacificus be promoted to the species level designation of Troglodytes pacificus with the common name of "Pacific wren". By applying a molecular clock to the amount of mitochondrial DNA sequence divergence between the two, it was estimated that Troglodytes pacificus and Troglodytes troglodytes last shared a common ancestor approximately 4.3 million years ago, long before the glacial cycles of the Pleistocene, which are thought to have promoted speciation in many avian systems inhabiting the boreal forest of North America. Two subspecies are accepted: * T. h. hiemalis Vieillot, 1819 – breeds in east Canada and northeast USA, winters in southeast USA * T. h. pullus (Burleigh, 1935) – breeds in mountains of West Virginia to Georgia (east-central USA), winters in south USA

The winter wren nests mostly in coniferous forests, especially those of spruce and fir, where it is often identified by its long and exuberant song. Although it is an insectivore, it can remain in moderately cold and even snowy climates by foraging for insects on substrates such as bark and fallen logs. Its movements as it creeps or climbs are incessant rather than rapid; its short flights swift and direct but not sustained, its tiny round wings whirring as it flies from bush to bush. At night, usually in winter, it often roosts, true to its scientific name, in dark retreats, snug holes and even old nests. In hard weather it may do so in parties, either consisting of the family or of many individuals gathered together for warmth.

===Breeding=== The male builds a small number of nests. These are called "cock nests" but are never lined until the female chooses one to use. The normal round nest of grass, moss, lichens or leaves is tucked into a hole in a wall, tree trunk, crack in a rock or corner of a building, but it is often built in bushes, overhanging boughs or the litter which accumulates in branches washed by floods. Five to eight white or slightly speckled eggs are laid in April, and second broods are reared.

Winter wrens primarily eat arthropods, with preference for spiders, caterpillars, and adult beetles and flies. Other less common food items include arthropods of taxa Acari (mites and ticks), Hymenoptera (bees, wasps, and ants), Chilopoda (centipedes), Diplopoda (millipedes), and Amphipoda (amphipods).